Moriguchi, C. Habutsu, N. Uno, H. (2026)A comparison of the ecological niches between the two sympatric ungulates, sika deer and Japanese serow, in a cool-temperate forest, central Japan.Mammal Research 71, 59(2026)➡
Seasonal Habitat Selection of Sika Deer in An Expanding Distribution Area, Northern Gunma Prefecture, Japan.Mammal Research 51, 3(2026)➡
Suzu Ogasawara, Yukiko Morimoto, Hiroyuki Uno(2026)Seasonal Habitat Selection of Sika Deer in An Expanding Distribution Area, Northern Gunma Prefecture, Japan.Mammal Study, 51(3):233-246 (2026). ➡
Takada H, Sun NC-M, Liang Y-J, Pei KJ-C. in press. Unique dietary partitioning among three sympatric forest-dwelling ungulates in Taiwan deviates from the Jarman–Bell principle. Contributions to Zoology DOI: 10.1163/18759866-bja10087
Takada H. Tezuka N. Yano R. Washida A. Hasegawa T. Yamazaki K. 2025. Competition between native ungulates: negative effects of sika deer (Cervus nippon) on foraging behavior, physiological stress, and population characteristics of Japanese serow (Capricornis crispus). Journal of Zoology 326: 403-413
Takada H. Watanabe H. Yano R. Tsukada H. 2025. Latrine ecology of a solitary ungulate, the Japanese serow: female-male communication site rather than territorial marking? Mammalia 89: 382-391
Selvarajah K. Baek SY. Takada H. Kim D. Yamada Y. Koike S. 2025. Variation in the activity patterns of Japanese serow (Capricornis crispus) across different temporal scales. European Journal of Wildlife Research 71: 136Nakamura K, Takada H, Koike S. 2025. Sex and age differences in habitat use priorities of sika deer (Cervus nippon) in highly seasonal habitats of Mount Fuji. Behaviour 165: 415-429.
Uno H, Iijima H. 2024. Annual variation in recruitment rates for the sika deer (Cervus nippon) population in eastern Hokkaido, Japan. Population Ecology 2024; 1-9.
Fukasawa k, Morosawa T, Nakashima Y, Takagi S, Yokoyama T, Ando M, Iijima H, Saito U M, Kumada N, Tochigi K, Yoshioka A, Funatsu S, Koike S, Uno H, Enomoto T, McShea W, Kays R. In press. Snapshot Japan 2023: the first camera trap dataset under a globally standardised protocol in Japan. Biodiversity data Journal.
Oishi K, Taniwaki T, Yasuda K, Dewi BS, Koike S (2026) Do exclosures in forests with high deer densities provide cover or food for rodents? Forest Ecology and Management
Selvarajah K, Baek S, Takada H, Kim D, Yamada Y, Koike S (2025) Variation in activity patterns of Japanese serow (Capricornis crispus) across different temporal scales. European Journal of Wildlife Research 71: 136.
Inagaki A, Allen M, Sebastian-Gonzalez E, Koike S (2025) Species roles and key interactions in vertebrate scavenger communities. Trends in Ecology and Evolution
Sawada S, Tochigi K, Kanamori H, Steyaert S, Koike S (2026) Variation in fat accumulation is associated with food availability in Asian black bears. Mammal study Nakamura K, Takada H, Koike S (2025) Sex and age differences in habitat use priorities of sika deer (Cervus nippon) in highly seasonal habitats of Mount Fuji. Behaviour
Inagaki A, Allen ML, Koike S (2025) Integration of specific metrics can promote the understanding of scavenging dynamics. Frontiers in Ecology and Evolution 13: 1611982
Koike S (2025) Variation in the Asiatic black bear Ursus thibetanus diet: a review. Mammal Study 50: 227-234.
Shimozuru M, Nakamura S, Yamazaki J, Yanagawa Y, Tamatani H,Kuroe M, Yamazaki K, Koike S, Goto Y, Naganuma T, Tochigi K, Inagaki A, Takekoshi N, Baek S, Sato N, Honda Y, Tsubota T, Ito H (2025) Epigenetic clock in bears: A simple cost-effective blood DNA methylation-based age estimation method applicable to multiple bear species. Ecology and Evolution 15: e71424.
Yamazaki K, Ishizuka M, Nakayama S, Banda N, Koike S, Goto Y, Yamazaki K, Nemoto Y, Fuchs B, Arnemo MJ (2025) Lead Concentration in Blood of Asian Black Bears in Two Areas of Central Japan. Science of the Total Environment 980: 179555.
Baek S, Zedrosser A, Yamazaki K, Goto Y, Takekoshi N, Koike S (2025) Risky behavior of Asian black bears differs between sex and season in a landscape fragmented by roads. Journal of Zoology /li>
Tsunamoto Y, Koike S, Tayasu I, Masaki T, Kato S, Kikuchi S, Nagamitsu T, Haraguchi T, Naganuma T, Naoe S (2025) Fruiting phenology affects the direction of vertical seed dispersal by mammals and birds across mountain ranges. Oecologia 207: 24.➡
Hayato Takada*, Akane Washida, Shuhei Yamasaki(2025)Ecological drivers of population abundance of the Japanese serow in the absence of predation risk.European Journal of Wildlife Research➡ プレスリリース
Seung-Yun Baek, Tatsuya Amano, Munemitsu Akasaka & Shinsuke Koike(2025) The range of large terrestrial mammals has expanded into human-dominated landscapes in Japan.Communications Earth & Environment➡ プレスリリース
Uno H, Iijima H. 2024. Annual variation in recruitment rates for the sika deer (Cervus nippon) population in eastern Hokkaido, Japan. Population Ecology 2024; 1-9.➡
Fukasawa k, Morosawa T, Nakashima Y, Takagi S, Yokoyama T, Ando M, Iijima H, Saito U M, Kumada N, Tochigi K, Yoshioka A, Funatsu S, Koike S, Uno H, Enomoto T, McShea W, Kays R. Snapshot Japan 2023: the first camera trap dataset under a globally standardised protocol in Japan. Biodiversity data Journal.➡
Dawson BM, Evans MJ, Barton PS, Soga M, Tochigi K, Koike S (2024) Drastic changes in ground-dwelling beetle communities following high-intensity deer culling: Insights from an island ecosystem. Environmental Entomology 53: 223-229.➡
Yasuda K, Taniwaki T, Amano T, Koike S (2024) Responses of a bird community to sporadic outbreaks of woody herbivorous insects in a temperate beech forest in Japan. Forest Ecology and Management 560: 121824.➡
Taniwaki T, Tamura A, Sashimura N, Naruse M, Tochigi K, Komine H, Koike S (2024) Indirect effects of low-impact thinning on insect communities in forest floor of coniferous plantations. Journal of Forest Research 29: 316-325.
Kazama K, Fujita K, Shinoda Y, Koike S (2024) Sika deer trajectory prediction considering environmental factors by timeseries transformer-based architecture. Expert Systems with Applications 250: 123630.➡
Maeda N, Baek S, Tochigi K, Naganuma T, Koike S (2024) Feeding habits of Japanese serow in a warm-temperate, low-elevation, evergreen broad-leaved shrub-dominated forest. Mammal Study 49: 273-287.
Takekoshi N, Fujitani A, Ohnishi N, Kozakai C, Koike S, Yamazaki K (2024) Factors affecting body weight fluctuation in free-ranging Asian black bears. Ursus35: e21.➡
Sato K, Goto Y, Koike S (2025) Epizoochorous seed dispersal by two Phasianidae birds: Green pheasant (Phasianus versicolor) and Chinese bamboo partridge (Bambusicola thoracicus). Ecological Research ➡
Inagaki A, Sugimoto Y, Allen ML, Koike S (2024) Documentation of an Asiatic black bear preying on a living sika deer caught in a leg-hold snare trap. Ursus➡ プレスリリース
Yasuda K, Kato D, Naoe S, Amanoa T, Yoshikawa T, Tochigi K, Koike S (2025) Does forest loss and fragmentation reduce woodpecker-associated ecosystem functions?. Global Ecology and Conservation➡ プレスリリース
Hiroyuki Uno, Hayato Iijima 2024 Annual variation in recruitment rates for the sika deer (Cervus nippon) population in eastern Hokkaido, Japan ➡
Hayato Takada*, Nick Ching-Min Sun, Yu-Jen Liang, Jen-Hao Liu, Ching-Kuo Liu, and Kurtis Jai-Chyi Pei 2024 Tree climbing behavior of a forest dwelling ungulate: The Formo-san serow.➡ プレスリリース
Hayato Takada*, Keita Nakamura 2024 Effects of human harvesting, residences, and forage abundance on deer spatial distribution.➡ プレスリリース
Hayato Takada*, Akiyoshi Sato, Setsuko Katsuta, Keita Nakamura, Atsushi Ohwaki, Yuto Kane, Takashi Hayakawa, Eiji Inoue 2024 Dietary partitioning in two gleaning bats, Japanese long-eared bats (Plecotus sacrimontis) and Ussuri tube-nosed bats (Murina ussuriensis): consuming insects of same order but different developmental stages.➡ プレスリリース
Kazuma Yasuda, Toru Taniwaki, Tatsuya Ama no, Shinsuke Koike 2024 Responses of a bird community to sporadic outbreaks of woody herbivorous insects in a temperate beech forest in Japan.Forest Ecology and Management.➡ プレスリリース
Hayato Takada*, Keita Nakamura, Masato Minami 2024 Diet selection of a solitary forest-dwelling ungulate, the Japanese serow (Capricornis crispus), in cool temperate forest.Mammal Study.➡ プレスリリース
Tezuka S, Tanaka M, Naganuma T, Tochigi K, Inagaki A, Myojo H, Yamazaki K, Allen M, Koike S. 2023. Comparing information derived on food habits of a terrestrial carnivore between animal-borne video systems and fecal analyses methods. Journal of Mammalogy 104: 184-193.➡
Takayama K, Ohnishi N, Zedrosser A, Anezaki T, Tochigi K, Inagaki A, Naganauma T, Yamazaki K, Koike S. 2023. Timing and distance of natal dispersal in Asian black bears. Journal of Mammalogy 104: 265-278.➡
Tanaka M, Baek S, Tochigi K, Naganuma T, Inagaki A, Dewi BS, Koike S. 2023. Conditions affecting ant nesting in stumps in a temperate coniferous planted forest. Forest Ecology and Management 537: 120976.
Baek S, Shimazaki A, Zedrosser A, Naganuma T, Kozakai C, Yamazaki K, Koike S. 2023. Response to human-modified landscape of an apex consumer: Sex- and season-related variations in road selection and movement pattern. Global Ecology and Conservation 46: e02603.➡
Inagaki A, Allen M, Maruyama T, Tochigi K, Koike S. 2023. Evidence for interspecific modulation of carcass consumption among facultative scavengers in Asian temperate forest. Food webs 37: e00324.
Takada H, Washida A, Yano R, Tezuka N, Minami M. 2023. Evolution from monogamy to polygyny: insights from the solitary Japanese serow. Behavioral Ecology and Sociobiology 77: 3➡
Hayato Takada*, Akiyoshi Sato, Setsuko Katsuta 2023.Seasonal diets of two rhinolophid bats Rhinolophus nippon and Rhinolophus cornutus in the Southern Japanese Alps.➡ プレスリリース
Kahoko Tochigi, Sam M.J.G. Steyaert, Keita Fukasawa, Misako Kuroe,Tomoko Anezaki, Tomoko Naganuma, Chinatsu Kozakai, Akino Inagaki, Koji Yamazaki, Shinsuke Koike 2023.Demographic Parameters of Asian Black Bears in Central Japan.Mammal Study.➡ プレスリリース
Takada H*, Nakamura K 2023 Overlap in habitat use and activity patterns between sika deer (Cervus nippon) and Japanese serows (Capricornis crispus) in subalpine habitats: exploitative competition rather than direct interference? Canadian Journal of Zoology.➡ プレスリリース
Takada H*, Yano R, Watanabe H, Ohuchi R, Kanno T, Washida A, Nakamura K, Tezuka N, Shimodoumae D, Minami M 2023 Behavioural interactions between sika deer and Japanese serows: are larger and gregarious ungulate dominant? Behaviour.➡ プレスリリース
Sato K, Goto Y, Koike S (2023) Seed attachment by epizoochory depends on animal fur, body height, and plant phenology. Acta Oecologia.➡ プレスリリース
Hiruma M, Takada H, Wasida A, Koike S. in press. Dietary partitioning and competition between sika deer and Japanese serows in high elevation habitats. Mammal Research. ➡ プレスリリース
Takada H. Unique spatial behavior of the Japanese serow (Capricornis crispus) in the open mountains of Mt. Fuji . Acta Ethologica.➡ プレスリリース
Hori M, Takada H, Nakane Y, Minami M, Inoue E. Genetic analysis reveals dispersal patterns of Japanese serow in two different habitats of a mountainous region. Zoological Science➡ プレスリリース
Morosawa, T. 2023. Synchronous decline of native bitterling population detected from 9 years survey in rivers and agricultural ditches around Lake Kasumigaura, Japan. Ichthyological Research 70:368–377.
Akashi N, Unno A, Uno H. 2022. The browsing ratio as an index of the impact of deer browsing on tree seedlings. Trees, Forests and People 8.➡
Allen M, Krofel M, Yamazaki K, Alexander EP, Koike S. 2022. Cannibalism in bears. Ursus 33: e10. ➡
Ando K, Yoshikawa T, Kozakai C, Yamazaki K, Naganuma T, Inagaki A, Koike S. 2022. Composite brownian walks best explain the movement patterns of Asian black bears, irrespective of sex, seasonality and food availability. Ecological Research 37: 522-531.➡
Hiruma M, Tochigi K, Kishimoto R, Kuroe M, Trentin EB, Koike S. 2022. Long-term stability in the winter diet of the Japanese serow. Zookeys 1122: 39-51.➡
Inagaki A, Allen M, Maruyama T, Yamazaki K, Tochigi K, Naganuma T, Koike S. 2022. Carcass detection and consumption by facultative scavengers in forest ecosystem highlights the value of their ecosystem services. Scientific Reports 12: 16451.➡ プレスリリース
Inatomi Y, Uno H, Ueno M, Takafumi H, Osa Y. 2022. Response of an indicator species, Dryopteris crassirhizoma, to temporal and spatial variations in sika deer density. Biology, 11, 302.➡
Jimbo M, Ishinazaka T, Shirane Y, Umemura Y, Yamanaka M, Uno H, Sashika M, Tsubota T, Shimozuru M. 2022. Diet selection and social learning: natal habitat influence on lifelong foraging strategies in solitary large mammals. Ecosphere 13:e4105.➡
Kawamura K, Jimbo M, Adachi K, Shirane Y, Nakanishi M, Umemura Y, Ishinazaka T, Uno H, Sashika M, Tsubota T, Shimozuru M. 2022. Diel and monthly activity pattern of brown bear and sika deer in the Shiretoko Peninsula, Hokkaido, Japan. The journal of Veterinary Medical Science 84(8): 1146-1156.">
Koike S, Tochigi K, Yamazaki K. 2023. Are seeds of trees with higher fruit production dispersed farther by frugivorous mammals? Journal of Forest Research 28: 64-72. ➡ プレスリリース
Naganuma T, Nakashita R, Tochigi K, Zedrosser A, Kozakai C, Yamazaki K, Koike S. 2022. Functional dietary response of Asian black bears to changes in sika deer density. Journal of Wildlife Management 86: e22218.➡
Oki K, Soga M, Amano T, Koike S. 2022. Abandonment of ski pistes impoverishes butterfly communities. Biodiversity and Conservation 31: 2083-2096.➡ プレスリリース
Osugi S, Baek S, Naganuma T, Tochigi K, Allen M, Koike S. 2022. The effect of decreasing human activity from COVID-19 on the foraging of fallen-fruit by omnivores. Ecology and Evolution 12: e9657.➡ プレスリリース
Osugi S, Trenti EB, Koike S. 2022. Effects of human activity on the fallen-fruit foraging behavior of Carnivora in an urban forest. Mammal Study 47: 113-123.➡
Takada H, Minami M. 2022. A preliminary study on habitat selection of the Japanese serow (Capricornis crispus) at two temporal scales, season and time of day, in a montane forest. Journal of Ethology 40: 91-95.➡
Takada H, Sato A, Katsuta S. 2022. Food habits of two species of tube-nosed bats, Murina hilgendorfi and Murina ussuriensis, in Hayakawa, central Japan. Animal Biology 72: 1-13.➡
Takada H, Washida A, Yano R, Tezuka N, Minami M. in press. Evolution from monogamy to polygyny: insights from the solitary Japanese serow. Behavioral Ecology and Sociobiology.➡ プレスリリース
Takayama K, Ohnishi N, Zedrosser A, Anezaki T, Tochigi K, Inagaki A, Naganauma T, Yamazaki K, Koike S. in press. Timing and distance of natal dispersal in Asian black bears. Journal of Mammalogy.➡ プレスリリース
Taniwaki T, Watanabe K, Komine H, Tochigi K, Yamane M, Koike S. 2022. Response of specialist and generalist predators to nonprogressive annual fluctuations in herbivorous insect population. Biological Control 165: 104810.➡
Tezuka S, Tanaka M, Naganuma T, Tochigi K, Inagaki A, Myojo H, Yamazaki K, Allen M, Koike S. in press. Comparing information derived on food habits of a terrestrial carnivore between animal-borne video systems and fecal analyses methods. Journal of Mammalogy.➡
Tochigi K, Steyaert S, Naganuma T, Yamazaki K, Koike S. 2022. Differentiation and seasonality in suitable microsites of seed dispersal by an assemblage of omnivorous mammals. Global Ecology and Conservation 40: e02335.➡ プレスリリース
Ito M, Takekoshi N, Koike S, Yamazaki K (2024) Den site utilization patterns of Japanese black bears during non-hibernation and hibernation periods. 17th Asian Society of Conservation Medicine Conference. Poster
Kushaal S, Baek S, Koike S (2024) Do serows wake up with the rising sun? International Society of Behavior Ecology. Poster
Koike S, Naganuma T, Baek S, Tochigi K, Takekoshi N, Yamazaki K (2024) What can we learn about bear ecology using animal-borne video systems? A case study of Asiatic black bears in Japan. 28th International Association for Bear Research and Management. Edmonton. Poster
Inagaki A, Allen ML, Tochigi K, Maruyama T, Koike S (2024) The role of Asiatic black bears Ursus thibetanus as scavenger in a Japanese temperate forest. 28th International Association for Bear Research and Management. Edmonton. Poster
Ohnishi N, Tochigi K, Naganuma T, Koike S, Yamazaki K, Osawa T (2024) Males disperse, females remain: revealing the dispersal patterns of Asian black bears through genetic analysis. 28th International Association for Bear Research and Management. Edmonton. Oral
Naganuma T, Baek S, Tochigi K, Takekoshi N, Kozakai C, Yamazaki K, Koike S (2024) Mating behavior and site selection of Asian black bear in Japan. 28th International Association for Bear Research and Management. Edmonton. Poster
Yamazaki K, Ishizuka M, Nakayama S, Banda N, Koike S, Goto Y, Yamazaki K, Nemoto Y, Fuchs B, Arnemo JM (2024) Japanese black bears are highly exposed to lead (Pb) . 28th International Association for Bear Research and Management. Edmonton. Poster
Makino T, Baek S, Inagaki A, Takekoshi N, Yamazaki K, Koike S (2024) Behavioral Characteristics of Autumn Migration in Solitary Terrestrial Mammals. 28th International Association for Bear Research and Management. Edmonton. Poster
Takekoshi N, Tomiyasu J, Yanagawa Y, Ohnishi N, Naganuma T, Baek S, Itoh M, Nishiwaki T, Kozakai C, Koike S, Yamazaki K (2024) Verification of the relationship between testosterone and behavioral changes in male Asian black bear (Ursus thibetanus). 28th International Association for Bear Research and Management. Edmonton. Poster
Baek S, Zedrosser A, Naganuma T, Yamazaki K, Koike S (2024) Asian black bear responses to roads: Sex- and season-related variations in road proximity, movement, and crossing. 28th International Association for Bear Research and Management. Edmonton. Oral
Uno H, Shimozuru M, Ishinazaka T, Yamanaka M, Fukazawa K.「Conservation and management of large mammals in the Shiretoko Natural World Heritage Site, Japan」（口頭発表）. 13th International Mammalogical Congress（第13回国際哺乳類学会）
Capture-recapture-recovery models for evaluating mortality,population density and animal space use in a heterogeneous landscape.2023. 13th International Mammalogical Congress (Anchorage, U.S.A.)
Inagaki A, Allen M, Naganuma T, Koike S (2022) Vertebrate Scavenging on Sika Deer Carcasses and Its Effects on Ecological Processes. Sika Deer: Life History Plasticity and Management 375-385. Springer.
Kaji K, Uno H, Iijima H. (2022) Future Challenges for Research and Management of Sika Deer. Sika Deer: Life History Plasticity and Management 615-634. Springer.
Koike S, Tochigi K, Soga M (2022) Impacts of Sika Deer Overabundance on the Structure and Functions of Dung Beetle Communities in Forest Ecosystems. Sika Deer: Life History Plasticity and Management 387-398. Springer.
Fujinami K, Naganuma T, Shinoda Y, Yamazak K, Koike S (2022) Attempts Toward Behavior Recognition of the Asian Black Bears Using an Accelerometer. Sensor- and Video-Based Activity and Behavior Computing 57-79, Springer.
Uno H, Inatomi Y, Ueno M, Kaji K. (2022) Adaptive Management of Sika Deer Populations on Hokkaido Island, Japan. Sika Deer: Life History Plasticity and Management 553-568. Springer.
メディア掲載情報
♦2026年度♦
2026年8月31日/The Asia Business Daily『Deer, Not Just Bears, Now Blamed for Landslides in Japan』/小池伸介教授
2026年8月28日/The Times of India『Japan killed more than 14,000 bears between April 2025 and January
2026 after record attacks on people; despite the cull, 6 people have been
killed this year and the government plans to remove another 8,800 bears』/小池伸介教授
